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Ventilation–Perfusion Matching in the Lung

1The V/Q Ratio as the Currency of Gas Exchange2Regional Heterogeneity: Gravity, Posture, and the Vertical V/Q Gradient3The Two Extremes: Shunt and Dead Space4Active Matching: Hypoxic Pulmonary Vasoconstriction and Bronchovascular Coupling
The V/Q Ratio as the Currency of Gas Exchange

Ventilation, Perfusion, and the Ratio That Binds Them

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Start with the two flows. Alveolar ventilation is the fresh air reaching the alveoli each minute, roughly four point two liters at rest. Perfusion is the mixed venous blood flowing through the pulmonary capillaries, roughly five liters, essentially the cardiac output. Divide them and you get a whole-lung ratio near zero point eight. That number is an average, not a description of any single alveolus. The reason the ratio matters more than either flow alone is that oxygen and carbon dioxide must cross between air and blood in the same alveolus. If air outruns blood, extra ventilation is wasted. If blood outruns air, blood leaves poorly oxygenated. The ideal single-alveolus model assumes every unit has the same ratio, which lets us predict gas pressures, but the real lung is a spread of ratios.
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Alveolar ventilation (V) and pulmonary perfusion (Q) are both flow rates measured in L/min. The whole-lung V/Q ratio is approximately 0.8 at rest, but this value is an average over a heterogeneous population of alveoli.

Why the ratio, not V or Q alone, determines gas exchange

Oxygen moves from alveolar gas into capillary blood, and carbon dioxide moves from blood into alveolar gas. For this exchange to be efficient, the flow of fresh air and the flow of mixed venous blood must arrive at the alveolus in the right proportion. If V is high and Q is low, a large volume of air contacts a small volume of blood, so the blood is well oxygenated but much of the ventilation is wasted. If Q is high and V is low, a large volume of blood contacts a small volume of air, so the blood leaves poorly oxygenated. The ratio V/Q captures this balance directly.

The ideal lung as a reference model

The ideal single-alveolar lung assumes every alveolus has the same V/Q ratio, equal to the whole-lung average. This model is useful for predicting how changes in V or Q shift alveolar and arterial partial pressures, but it does not describe the real lung, where V/Q varies from region to region. The real lung is a spectrum of ratios, and that heterogeneity has its own consequences for arterial oxygenation.

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